Bulletin Mar 2003: Arm in Arm. The capsid, or outer coat, of the papillomavirus is made of 72 ring-like structures. Each is connected to its neighbors via a type of outstretched arm arrangement. Harrison and colleagues believe this assembly affects the way the virus appears to the immune system. / Annual Report 2004: Micromanagement. Stephen Harrison’s laboratory studies the atomic structures of macromolecular assemblies, such as in this papilloma virus, to understand how they function in cells.